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Aylward CM, Grahn R, Barthman-Thompson L, Kelt D, Sacks BN, Statham MJ. A Novel Non-Invasive Genetic Survey Technique for Small Mammals. Journal of Mammalogy. June 2022.

Aylward CM, Statham MJ, Barthman-Thompson L, Kelt D, Sacks BN. Dietary characterization of the Endangered Salt Marsh Harvest Mouse and Sympatric Rodents using DNA Metabarcoding. Ecology and Evolution. July 2022.

Statham MJ, Aylward CM, Barthman-Thompson L, Kierepka EM, Sacks BN. Landscape genetics of an endangered salt marsh endemic: identifying population continuity and barriers to dispersal. Conservation Genetics.


McDevitt AD, Coscia I, Browett SS, Ruiz- González A, Statham MJ, et al. (2021) Next-generation phylogeography resolves post-glacial colonization patterns in a widespread carnivore, the red fox (Vulpes vulpes), in Europe. Molecular Ecology, Nov. 2021.

Statham MJ, Barthman-Thompson L, Fresquez S, Sacks BN (2021). Development of a morphological key for the southern salt marsh harvest mouse using genetically verified individuals. California Fish and Wildlife, CESA Special Issue 2021. Cover article.

Supporting information for the California Fish and Wildlife Journal. Harvest mouse ventral coloration photographs.

Cover of special issue on California Endangered Species

Sacks BN, Mitchell KJ, Quinn CB, Hennelly LM, Sinding MHS, Statham MJ et al. (2021) Pleistocene origins, western ghost lineages, and the emerging phylogeographic history of the red wolf and coyote. Molecular Ecology. 17: 4292-4304. Cover article.

Alminas OSV, Heffelfinger JR, Statham MJ, Latch EK. (2021) Phylogeography of Cedros and Tiburón Island mule deer in North America’s desert southwest. Journal of Heredity. 112: 260–275


Castle ST, Allan N, Clifford D, Aylward CM, Ramsey J, Fascetti A, Pesapane R, Roy A, Statham MJ, Sacks BN, Foley J. (2020) Diet composition analysis provides new management insights for a highly specialized endangered small mammal. PLoS ONE 15(10): e0240136.


Statham MJ, Woollett (Smith) D, Fresquez S, Pfeiffer JU, Richmond J, Whitelaw A, Richards N, Westphal MF, Sacks BN. (2019) Noninvasive identification of herpetofauna: pairing conservation dogs and genetic analysis. Journal of Wildlife Management. 84: 66-74.

U = Undergraduate Student

Alexander N, Statham MJ, Sacks BN, Bean WT. (2019) Generalist dispersal and gene flow of an endangered keystone habitat specialist (Dipodomys ingens). Journal of Mammalogy. 100: 1533-1545.

Statham MJ, Bean WT, Westphal M, Alexander N, Sacks BN. (2019) Historical population size change and differentiation of relict populations of the endangered giant kangaroo rat. Journal of Heredity. 110: 548-558.


Statham MJ, Edwards CJ, Norén K, Soulsbury CD, Sacks BN (2018). Genetic analysis of European red foxes reveals multiple distinct peripheral populations and central continental admixture. Quaternary Science Reviews. 197: 257-266.

Smith KR, Riley MK, Barthman-Thompson L, Woo I, Statham MJ, Estrella S, Kelt DA (2018). Toward Salt Marsh Harvest Mouse Recovery: A Review. San Francisco Estuary and Watershed Science. 16: 1-24?.

Smith KR, Riley MK, Barthman-Thompson L, Statham MJ, Estrella S, Kelt DA (2018). Toward Salt Marsh Harvest Mouse Recovery: Research Priorities. San Francisco Estuary and Watershed Science. 16: 1-6?.

Sacks BN, Statham MJ, Lounsberry Z (2018). Nuclear genetic analysis of the red fox across its trans-Pacific range. Journal of Heredity.

Wallén J, Statham, MJ, Ågren E, Isomursu M, Flagstad O, Bjorneboe Berg T, Sacks B, Noren K. Multiple recolonization routes towards the north: population history of the Fennoscandian red fox (Vulpes vulpes). Biological Journal of the Linnean Society.


Yannic G, Statham MJ, Denoyelle L, Szor G, Qulaut GQ, Sacks BN, Lecomte N (2017). Investigating the ancestry of putative hybrids: are Arctic fox and red fox hybridizing? Polar Biology, 40: 2055-2062.

Merson C, Statham MJ, Janecka JE, Lopez RR, Silvy NJ, Sacks BN (2017). Distribution of native and nonnative ancestry in red foxes along an elevational gradient in central Colorado. Journal of Mammalogy, 98: 365-377.
Lounsberry ZT, Quinn CB, Statham MJ, Angulo CLU, Kalani TJU, Tiller E, Sacks BN (2017). Investigating genetic introgression from farmed red foxes into the wild population in Newfoundland, Canada. Conservation Genetics, 18: 383-392.


Statham M, Aamoth SU, Barthman-Thompson L, Estrella S, Fresquez S, Hernandez L et al. (2016) Conservation genetics of the endangered San Francisco Bay endemic salt marsh harvest mouse (Reithrodontomys raviventris). Conservation Genetics, 17: 1055-1066

Sacks BN, Statham MJ, Wittmer H (2016). A preliminary range-wide distribution model for the Sacramento Valley red fox (Vulpes vulpes patwin). Journal of Fish and Wildlife Management. 8: 1-11.

Gulsby W, Killmaster C, Bowers J, Laufenberg J, Sacks B, Statham M, Miller K (2016). Efficacy and precision of fecal genotyping to estimate coyote abundance in an area with a humid subtropical climate. Wildlife Society Bulletin. Accepted July 2016.


Kasprowicz A, Statham MJ, Sack BN. (2015) The fate of the other red coat: remnants of colonial British red foxes in the Eastern United States. Journal of Mammalogy.

Goddard NU, Statham MJ, Sacks BN (2015). Mitochondrial analysis of the most basal canid reveals deep divergence between eastern and western North American gray foxes (Urocyon spp.) and ancient roots in Pleistocene California. PLoS ONE. 10(8): e0136329. oi:10.1371/journal.pone.0136329

U = undergraduate author

Norén K, Statham MJ, Ågren E, Isomursu M, Flagstad Ø, Eide NE, et al. (2015) Tracking genetic footprints to reconstruct geographic patterns of expansion in Fennoscandian red foxes. Global Change Biology.

Volkmann LA, Statham MJ, Mooers AØ, Sacks BN. (2015) Genetic distinctiveness of red foxes in the Intermountain West as revealed through expanded mitochondrial sequencing. Journal of Mammalogy. 96: 297-307.

Gulsby WD, Killmaster CH, Bowers JW, Kelly JD, Sacks BN, Statham MJ, et al. (2015) White-tailed deer fawn recruitment before and after experimental coyote removals in central Georgia. Wildlife Society Bulletin.

Berteaux D, Gallant D, Sacks BN, Statham MJ. (2015) Red foxes (Vulpes vulpes) at their expanding front in the Canadian Arctic have indigenous maternal ancestry. Polar Biology.


Statham MJ*, Murdoch J, Janecka J, Aubry KB, Edwards JE, Soulsbury CD, et al. (2014) Range-wide multilocus phylogeography of the red fox reveals ancient continental divergence, minimal genomic exchange, and distinct demographic histories. Molecular Ecology, 23: 4813–4830. Altmetrics score = 64

Reponen SEM, Statham MJ*, Barthman-Thompson L, Sacks BN (2014) Microsatellite primer development for the salt marsh harvest mouse (Reithrodontomys raviventris) and cross-amplification in the western harvest mouse (R. megalotis). Conservation Genetics Resources, 6: 285-287.


Statham MJ*, Sacks BN, Aubry KB, Perrine JD, Wisely SM (2012) The origin of recently established red fox populations in the contiguous United States: translocations or natural range expansions? Journal of Mammalogy, 93(1): 52-65.

Edwards CJ, Soulsbury C, Statham MJ, Ho S, et al. (2012) Temporal genetic variation of the red fox, Vulpes vulpes, across western Europe and the British Isles. Quaternary ScienceReview, 57: 95-104.

Statham MJ*, Rich AR, Lisius SK, Sacks BN (2012) Discovery of a remnant population of Sierra Nevada red fox (Vulpes vulpes necator). Northwest Science, 86(2): 122-132. Featured article on BioOne Webpage.

O’Meara DB, Edwards CJ, Sleeman DP, McDowell JR, Coughlan JP, Dillane E, Cross TF, O’Leary D, Statham MJ, Pope LC, O’Reilly C, Bradley DG, Carlsson J (2012) Mitochondrial and microsatellite DNA analyses of the Eurasian badgers (Meles meles) in Ireland and Britain, compared to Western Europe. Biological Journal of the Linnean Society, 106: 893-909.


Sacks BN, Moore M, Statham MJ, Wittmer HU (2011) A restricted hybrid zone between native and introduced red fox (Vulpes vulpes) populations suggests reproductive barriers. Molecular Ecology, 20: 326-341.


Statham MJ*, Trut LN, Sacks BN, Kharlamova AV, Oskina IN, Gulevich RG, Johnson JL, Temnykh SV, Acland GM, Kukekova, AV (2011) On the origin of a domesticated species: identifying the parent population of Russian silver foxes (Vulpes vulpes). Biological Journal of the Linnean Society, 103: 168–175.


Sacks BN, Statham MJ, Perrine JD, Wisely SM, Aubry KB (2010) North American montane red foxes: expansion, fragmentation, and the origin of the Sacramento Valley red fox. Conservation Genetics, 11: 1523-39.

Mullins J, Statham MJ, Roche T, Turner PD, O’Reilly C (2010) Remotely plucked hair genotyping: a reliable and non-invasive method for censusing pine marten (Martes martes, L. 1758) populations. European Journal of Wildlife Research, 56(3): 443-453.



Sacks BN, Våge DI, Statham MJ (2009) A medium-throughput SNP assay for detecting genetic variation in coding and non-coding portions of the red fox genome. Conservation Genetics Resources, 1: 459–463.

Conard JM, Statham MJ, Gipson PS, Wisely SM (2009) The influence of translocation strategy and management practices on the genetic variability of a re-established elk (Cervus elaphus) population. Restoration Ecology, 18: 85-93.

Aubry KB, Statham MJ, Sacks BN, Perrine JP, Wisely SM (2009) Phylogeography of the North American red fox: vicariance in Pleistocene forest refugia. Molecular Ecology, 18: 2668-2686. Cover article.


O’Reilly C, Statham M, Mullins J, Turner PD, O’Mahony D (2008) Efficient species identification of pine marten (Martes martes) and red fox (Vulpes vulpes) scats using a 5′ nuclease real-time PCR assay. Conservation Genetics, 9: 735-738.

Wisely SM, Statham MJ, Fleischer RC (2008) Pleistocene refugia and Holocene expansion of a grassland dependent species, the black-footed ferret (Mustela nigripes). Journal of Mammalogy, 89(1): 87-96.

Statham MJ*, Turner PD, O’Reilly C (2007) Molecular Sex Identification of five Mustelid species. Zoological Studies, 46(5): 600-608.


Sleeman DP, Statham M, Dixon C (2006) Records of Otters (Lutra lutra L.) and Badgers (Meles meles L.) from built-up areas in Ireland. Bulletin of the Irish Biogeographical Society, 30: 3-6.


Statham M, Turner PD, O’Reilly C (2005) Use of PCR amplification and restriction enzyme digestion of mitochondrial D-loop for identification of mustelids in Ireland. Irish Naturalist’s Journal, 28: 1-6.


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